Orthoptera Species FileOrthoptera Species File

species

Conocephalus (Anisoptera) maculatus (Le Guillou, 1841)

Spotted Meadow Katydid

Conocephalus (Anisoptera) maculatus (Le Guillou, 1841): male (Siargao Island, 16 October 2018). (Otu).
Conocephalus (Anisoptera) maculatus (Le Guillou, 1841): male (Siargao Island, 16 October 2018). (Otu).
Conocephalus (Anisoptera) maculatus (Le Guillou, 1841): Fig. 8. male last abdominal tergite (Conocephalus bidens). (Otu).
Conocephalus (Anisoptera) maculatus (Le Guillou, 1841): Fig. 9. male cercus (Conocephalus bidens). (Otu).
CollectionObject 1516767; 90e7eec5-475f-448a-b878-5d2404dc232e: male, lateral view (neotype). (CollectionObject).
CollectionObject 1516767; 90e7eec5-475f-448a-b878-5d2404dc232e: stridulatory area (neotype). (CollectionObject).
CollectionObject 1516767; 90e7eec5-475f-448a-b878-5d2404dc232e: male cercus (neotype). (CollectionObject).
CollectionObject 1516772; 39469363-e272-4d39-86e9-8ecc99b5c659: male, lateral view (holotype de Conocephalus arabicus). (CollectionObject).
CollectionObject 1516772; 39469363-e272-4d39-86e9-8ecc99b5c659: male, lateral view (holotype de Conocephalus arabicus). (CollectionObject).
CollectionObject 1516772; 39469363-e272-4d39-86e9-8ecc99b5c659: male head and pronotum, lateral view (holotype de Conocephalus arabicus). (CollectionObject).
CollectionObject 1516772; 39469363-e272-4d39-86e9-8ecc99b5c659: male, dorsal view (holotype de Conocephalus arabicus). (CollectionObject).
CollectionObject 1516772; 39469363-e272-4d39-86e9-8ecc99b5c659: male pronotum and stridulatory apparatus (holotype de Conocephalus arabicus). (CollectionObject).
CollectionObject 1516772; 39469363-e272-4d39-86e9-8ecc99b5c659: male cerci (holotype de Conocephalus arabicus). (CollectionObject).
CollectionObject 1516772; 39469363-e272-4d39-86e9-8ecc99b5c659: male cerci (holotype de Conocephalus arabicus). (CollectionObject).
CollectionObject 1516772; 39469363-e272-4d39-86e9-8ecc99b5c659: labels (holotype de Conocephalus arabicus). (CollectionObject).
CollectionObject 1516773; 3c6b61b2-ea2f-4621-a104-9b0d0849570f: male, lateral view (holotype of Conocephalus bidens). (CollectionObject).
CollectionObject 1516773; 3c6b61b2-ea2f-4621-a104-9b0d0849570f: male head and pronotum, lateral view (holotype of Conocephalus bidens). (CollectionObject).
CollectionObject 1516773; 3c6b61b2-ea2f-4621-a104-9b0d0849570f: male, dorsal view (holotype of Conocephalus bidens). (CollectionObject).
CollectionObject 1516773; 3c6b61b2-ea2f-4621-a104-9b0d0849570f: male stridulatory area (holotype of Conocephalus bidens). (CollectionObject).
CollectionObject 1516773; 3c6b61b2-ea2f-4621-a104-9b0d0849570f: male cerci (holotype of Conocephalus bidens). (CollectionObject).
CollectionObject 1516773; 3c6b61b2-ea2f-4621-a104-9b0d0849570f: labels (holotype of Conocephalus bidens). (CollectionObject).
CollectionObject 1522134; dfb531f0-8d4f-4df2-9b23-8a34f90db542: male, lateral view (holotype of Xiphidium neglectum). (CollectionObject).
CollectionObject 1586780; f183771f-fa12-49f3-a1e0-243837c986ca: male, lateral view, scale bar = 5 mm. (CollectionObject).

Type specimen

Neotype; male; 90e7eec5-475f-448a-b878-5d2404dc232e; deposited at: Natural History Museum, London (NHMUK); West Malaysia

Nomenclature (97)

  • Xiphidion maculatus Le Guillou, 1841: 294.

    Neotype; male; 90e7eec5-475f-448a-b878-5d2404dc232e; deposited at: Natural History Museum, London (NHMUK); West Malaysia

  • Locusta (Xiphidium [sic]) lepida Haan, 1843: 188, 189.

    Syntype; male; 96d9df3f-4a0f-4ca5-af17-c33322071755; deposited at: Naturalis Biodiversity Center (NBC);

  • ... Show all ... (93)
  • Conocephalus (Anisoptera) maculatus (Le Guillou, 1841) in Massa, 2025: 17.
  • Conocephalus (Anisoptera) maculates [sic] Le Guillou, 1841 (misspelling of Xiphidion maculatus Le Guillou, 1841)

Nomenclature references (76)

  • Bey-Bienko, G.Y. (1957) Results of Chinese-Soviet zoological-botanical expeditions to south-western China 1955-1956: Tettigonioidea (Orthoptera) of Yunnan. Entomologicheskoe Obozrenie, 36, 401–417.
  • Bruner, L. (1915) Preliminary catalogue of the Orthopteroid insects of the Philippine Islands. University Studies, 15(2), 195–281 [1–87]. Available at http://www.biodiversitylibrary.org/item/94891#page/233/mode/1up
  • ... Show all ... (72)
  • Bruner, L. (1920) Orthoptera from Africa, being a report upon some Saltatoria mainly from Cameroon contained in the Carnegie Museum. Annals of the Carnegie Museum, 13, 92–142. [1919] Available at http://www.biodiversitylibrary.org/item/38118
  • Brunner von Wattenwyl, C. (1893) Révision du système des Orthoptères et déscription des espèces rapportées par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, 13(33), 1–230. Available at http://www.biodiversitylibrary.org/item/25035
  • Chopard, L. (1954) La Réserve naturelle intégrale du Mt. Nimba, III. Orthoptères Ensifères. Mémoires de l'Institut Français d'Afrique Noire, 40(2), 25–97.
  • Dharini, S.V. & Chitra, N. (2023) Diagnostics of the tettigoniid genus Conocephalus occurring in the rice fields. Indian Journal of Entomology, 85(2), 359–366. https://doi.org/10.55446/ije.2022.434
  • Ebner, R. (1939) Tettigoniiden (Orthoptera) aus China. Lingnan Science Journal, 18, 293–302, 11 figs.
  • El-Hawagry, M.A., Khalil, M.W., Sharaf, M.R., Fadl, H.H. & Aldawood, A.S. (2013) A preliminary study on the insect fauna of Al-Baha Province, Saudi Arabia, with descriptions of two new species. ZooKeys, 274, 1–88. Available at http://www.pensoft.net/journal_home_page.php?journal_id=1&page=article&SESID=&type=show&article_id=4529
  • Farooqi, M.K. & Usmani, M.K. (2018) Review of genus Conocephalus Thunberg, 1815 (Orthoptera: Tettigoniidae: Conocephalinae) with one new species from India. Zootaxa, 4461(3), 381–398.
  • Farooqi, M.K., Ali, M., Amir, M. & Usmani, M.K. (2022) First description of the male Conocephalus (Anisoptera) hilli Farooqi & Usmani, 2019 in rice fields from India. Transactions of the American Entomological Society, 148(2), 187–195.
  • Gaikwad, S.M., Koli, Y.J., Raut, G.A., Waghmare, S.H. & Bhawane, G.P. (2016) Long-horned grasshoppers (Orthoptera: Tettigoniidae) in Radhanagari Wildlife Sanctuary, Maharashtra, India. Journal of Threatened Taxa, 8(2), 8533–8537. Available at http://www.threatenedtaxa.in/index.php/JoTT/article/view/2574
  • Govindaraj, D. & Natchiappan, S. (2018) Influence of courtship feeding on fecundity of Conocephalus maculatus (Le Guillou, 1841) (Orthoptera: Tettigoniidae: Conocephalinae). Journal of Entomology and Zoology Studies, 6(5), 612–617. Available at http://www.entomoljournal.com/archives/2018/vol6issue5/PartJ/6-4-301-584.pdf
  • Griffini, A. (1906) Ortotteri raccolti da Leonardo Fea nell'Africa occidentale. 1. Hetrodidi, Conocephalidi, Meconemidi, Pseudophyllidi, Mecopodidi e Fanerotteridi. Annali del Museo Civico di Storia Naturale di Genova, 3(2), 358–397. Available at http://www.biodiversitylibrary.org/item/137853#page/366/mode/1up
  • Griffini, A. (1908) Phasgonuridae africane del R. Museo di Storia Naturale di Bruxelles. Meconemidae et Conocephalidae. Mémoires de la Société entomologique de Belgique, 15, 65–73. Available at http://www.biodiversitylibrary.org/item/41740#page/79/mode/1up
  • Hebard, M. (1922) Studies in Malayan, Melanesian and Australian Tettigoniidae (Orthoptera). Proceedings of the Academy of Natural Sciences of Philadelphia, 74, 121–299, pl. 11–22. Available at http://www.archive.org/details/proceedingsofaca74acad
  • Heller, K.-G. (2019) Provisional checklist of the Tettigonioidea (Insecta: Orthoptera) from São Tomé & Príncipe with taxonomic remarks, bioacoustical data and the description of new taxa. Zootaxa, 4563(1), 41–66.
  • Hemp, C. & Hemp, A. (2003) Saltatoria coenoses of high-altitude grasslands on Mt. Kilimanjaro Tanzania (Orthoptera: Saltatoria). Ecotropica, 9, 71–97. Available at http://www.soctropecol.eu/publications/pdf/9%201-2/Hemp,%20Hemp,%202003.pdf
  • Hemp, C. (2013) Annotated list of Ensifera (Orthoptera) and further records on Caelifera (Orthoptera) of Mt Kilimanjaro, Tanzania. Zootaxa, 3613(4), 301–342.
  • Hemp, C. (2021) In A Field Guide to the Bushcrickets, Wetas and Raspy Crickets of Tanzania and Kenya. Senckenberg. 451 pp.
  • Henry, G.M.R. (1932) Notes on Ceylon Tettigoniidae, with descriptions of new species. Part 1. Ceylon Journal of Science Biological Science), 16(3), 229–256.
  • Ingrisch, S. & Garai, A. (2001) Orthopteroid Insects from Ganesh Himal, Nepal. Esperiana, Schwanfeld, 8, 755–770.
  • Ingrisch, S. & Shishodia, M.S. (1998) New species and records of Tettigoniidae from India (Ensifera). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 71(3-4), 355–371. Available at http://www.e-periodica.ch/digbib/view?pid=seg-001%3A1998%3A71%3A%3A275#372
  • Ingrisch, S. & Shishodia, M.S. (2000) New taxa and distribution records of Tettigoniidae from India (Orthoptera: Ensifera). Mitteilungen der Münchner Entomologischen Gesellschaft, 90, 5–37. Available at http://www.biodiversitylibrary.org/page/29007178#page/135/mode/1up
  • Ingrisch, S. (1990) Grylloptera and Orthoptera s.str. from Nepal and Darjeeling in the Zoologische Staatssammlung München. Spixiana (Munich), 13, 149–182. Available at http://archive.org/details/spixiana1990zool
  • Ingrisch, S. (1990) Zur Laubheuschrecken-Fauna von Thailand (Insecta: Saltatoria: Tettigoniidae). Senckenbergiana biologica, 70(1-3), 89–138. [1989]
  • Ingrisch, S. (1999) Orthopteroid insects of Yemen. Esperiana, Schwanfeld, 7, 349–376.
  • Ingrisch, S. (2002) Orthoptera from Bhutan, Nepal and North India in the Natural History Museum Basel. Entomologica Basiliensia, 24, 123–159. Available at http://s.ingrisch.macbay.de/EN/page2/page2.html
  • Ito, G. & Ichikawa, A. (2004) Notes on Matsumura's type specimens of Orthoptera. Insecta Matsumurana, New Series, 60, 55–65. [2003] Available at http://eprints.lib.hokudai.ac.jp/dspace/bitstream/2115/9918/1/60_p55-65.pdf
  • Jacobson, G.G. & Bianchi, V.L. (1905) In Orthopteroid and Pseudoneuropteroid Insects of Russian Empire and adjacent countries. Devrien Publ., St. Petersburg. i–xii, 1–952, pls 1–25. [1902]
  • Jin, X. & Xia, K.-L. (1994) An index-catalogue of Chinese Tettigoniodea (Orthopteroidea: Grylloptera). Journal of Orthoptera Research, 3, 15–41.
  • Karny, H.H. (1907) Revisio Conocephalidarum. Abhandlungen der K.K. Zoologisch-botanischen Gesellschaft Wien, 4(3), 1–114. Available at http://www.landesmuseum.at/biophp/de/band_det.php?litnr=29695
  • Karny, H.H. (1912) Orthoptera Fam. Locustidae Subfam. Conocephalinae. In Genera Insectorum. P. Wytsman [Ed.]. V. Verteneuil & L. Desmet, Bruxelles. Vol. 135, 1–17 pp. Available at http://www.biodiversitylibrary.org/item/123362#page/3/mode/1up
  • Karny, H.H. (1915) H. Sauter's Formosa-Ausbeute. Orthoptera et Oothecaria. Supplementa Entomologica, 4, 56–108. Available at http://sdei.senckenberg.de/~openaccess/01469.pdf
  • Karny, H.H. (1923) On Malaysian katydids (Gryllacridae and Tettigoniidae) from the Raffles Museum, Singapore. Journal of the Malaysian Branch of the Royal Asiatic Society, 1, 117–193.
  • Karny, H.H. (1926) Fauna Buruana. Orthoptera, Fam. Tettigoniidae. Hortus Botanicus Bogoriensis, Java (Treubia), 7, 145–215. [1925] Available at http://e-journal.biologi.lipi.go.id/index.php/treubia/article/view/1717
  • Kim, T.-W. & Kim, J.-I. (2002) Taxonomic study of the genus Conocephalus Thunberg in Korea (Orthoptera: Tettigoniidae: Conocephalinae). Korean Journal of Entomology, 32(1), 13–19. Available at http://jasa.pe.kr/pulmuchi/conocephalus.pdf
  • Kirby, W.F. (1906) Orthoptera Saltatoria. Part I. (Achetidae et Phasgonuridae). In A Synonymic Catalogue of Orthoptera (Orthoptera Saltatoria, Locustidae vel Acridiidae). British Museum (Natural History), London. Vol. 2, i-viii, 1–562. Available at http://www.biodiversitylibrary.org/item/29662
  • Le Guillou, E.J.F. (1841) Description des orthoptères nouveaux, recueillis pendant son voyage de circumnavigation sur la corvette la Zélée. Revue et Magasin de Zoologie Pure et Appliquée, 291–295.
  • Lee, S.M. (1990) Systematic notes on Tettigoniidae of Korea. INSECTA KOREANA, 7, 104–117.
  • Li, Y. & Shi, F. (2018) Notes on the genus Conocephalus Thunberg, 1815 (Orthoptera: Tettigoniidae: Conocephalinae) in Southwest China with description of one new species. Zootaxa, 4438(1), 148–158.
  • Liu, X.-W. & Zhang, D. (2007) A study of the genus Conocephalus Thunberg and descriptions of two new species from China (Orthoptera, Conocephalidae). Acta Zootaxonomica Sinica, 32, 438–444.
  • Mahasneh, A. & Katbeh-Bader, A. (2004) A taxonomic study on the long-horned grasshoppers of Jordan (Orthoptera: Tettigoniidae). Denisia 04, zugleich Kataloge des Oberösterreichischen Landesmuseums, Neue Folge, 14, 245–264.
  • Massa, B. (2009) Annotated check-list of Orthoptera of Libya. Journal of Orthoptera Research, 18(1), 75–93. Available at http://www.bioone.org/doi/full/10.1665/034.018.0109
  • Massa, B. (2021) Tettigoniidae (Insecta: Orthoptera) collected in tropical forests of Zambia, Cameroon, Gabon and São Tomé during the entomological expeditions of African Natural History Research Trust. Annales de la Société Entomologique de France, 57(1), 29–76.
  • Massa, B. (2025) Diversity of Leaf Katydids of Odzala-Kokoua National Park, Republic of Congo, Central Tropical Africa (Insecta: Orthoptera: Tettigoniidae). Insects, 16(3: 241), 1–71. https://doi.org/10.3390/insects16030241
  • Matsumura, S. & Shiraki, T. (1908) Locustiden Japans. Journal of the College of Agriculture, Tohoku Imperial University, 3(1), 1–80.
  • Nagar, R. & Swaminathan, R. (2016) Notes on the genus Conocephalus (Orthoptera: Tettigoniidae; Conocephalinae; Conocephalini) from India and description of two new species. Zootaxa, 4126(1), 1–43.
  • Naskrecki, P. & Guta, R. (2019) Katydids (Orthoptera: Tettigoniidae) of Gorongosa National Park and Central Mozambique. Zootaxa, 4682(1), 1–119.
  • Oda, K. & Ishii, M. (1998) Factors affecting color polymorphism in the meadow grasshopper, Conocephalus maculatus (Orthoptera: Tettigoniidae). Applied Entomology and Zoology, 33, 455–460.
  • Otte, D. (1997) Orthoptera Species File 7. Tettigonioidea. The Orthopterists' Society and The Academy of Natural Sciences of Philadelphia, Philadelphia, 373 pp.
  • Panhwar, W.A., Sultana, R., Wagan, M.S. & Kumar, S. (2013) On the distribution and taxonomy of Conocephalus species (Orthoptera: Tettigonioidea: Conocephalinae) from Pakistan. Journal of Biodiversity and Environmental Sciences (JEBS), 3(11), 171–176. Available at http://www.innspub.net/wp-content/uploads/2013/10/JBES-Vol3No11-p171-176.pdf
  • Pitkin, L.M. (1980) A revision of the Pacific species of Conocephalus Thunberg (Orthoptera: Tettigoniidae). Bulletin of the British Museum (Natural History) Entomology, 41(5), 315–355. Available at http://www.archive.org/details/bulletinofbritis41entolond
  • Popov, G.B. (1981) Orthoptera: Family Tettigoniidae (Bush Crickets). In Fauna of Saudi Arabia. Ciba-Geigy Ltd., Basel. Vol. 3, pp. 114–148.
  • Ragge, D.R. & Roy, R. (1961) Le Parc National du Niokolo-koba VII. Orthoptera Tettigonioidea. Mémoires de l'Institut Français d'Afrique Noire, 62, 105–107.
  • Redtenbacher, J. (1891) Monographie der Conocephaliden. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 41, 315–562. Available at http://www.biodiversitylibrary.org/item/49919#page/459/mode/1up
  • Roy-Noel, J. (1954) Contribution à l'étude de l'appareil musical des orthoptères ensifères. Annales des Sciences Naturelles, 16, 65–83.
  • Shi, F. & Wang, P. (2015) The genus Conocephalus Thunberg (Orthoptera: Tettigoniidae: Conocephalinae) in Hainan, China with description of one new species. Zootaxa, 3994(1), 142–144.
  • Shishodia, M.S. (2006) Insecta: Orthoptera: Tettigoniidae. In Fauna of Nagaland. Zoological Survey of India, Calcutta. 89–93 pp. Available at http://faunaofindia.nic.in/PDFVolumes/sfs/053/index.pdf
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© Basemap from GBIF

Descendants and synonyms

  • Conocephalus (Anisoptera) maculatus (Le Guillou, 1841)
    • =Conocephalus (Xiphidion) arabicus Uvarov, 1933
    • =Conocephalus bidens Uvarov, 1957
    • =Xiphidium [sic] continuum Walker, 1869
    • =Xiphidium [sic] dimidiatum Matsumura & Shiraki, 1908
    • =Locusta (Xiphidium [sic]) lepida Haan, 1843
    • =Conocephalus (Anisoptera) maculates [sic] Le Guillou, 1841
    • =Xiphidion neglectum Bruner, 1920
Life zone and ecology data

Terrestrial. Ubiquitous in habitats with lowland grass, including roadside verges and pastures.

Sound links
External links to websites
Gender, form, and etymology
Adjectivemaculatus, maculata, maculatum
Conocephalus (Anisoptera) maculatus (Le Guillou, 1841)

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